Glossary
Page Last Updated: January 18 2021
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Phyllaries - bracts at the base of a composite (Aster family) flower head - also known as involucral
bracts - the picture at right shows phyllaries of Tall Beggar-ticks - the picture at far right shows the
phyllaries of Knapweed
Cultivar - a 'cultivated variety" of a plant species that has been bred for desirable characteristics
such as flower colour or shape, crop yield or resistance to disease - genetically, a cultivar is not
sufficiently different from its host to be defined as a new species - the picture at right shows the
'Crimson King' cultivar of the Norway Maple tree
Ootheca - an egg mass with a protective outer covering - produced by certain insects (notably
mantids and cockroaches) and mollusks - usually contains many eggs - the picture at right shows
the ootheca of a Praying Mantis
Autoecious - Rust fungus that can complete its life cycle on one host species - Heteroecious -
Rust fungus that requires two unrelated host species to complete its life cycle - there may be many
different potential hosts for a given species of Rust fungus - the picture at right shows the
Heteroecious Crown Oat Rust (hosts are Grass (barley, rye) and Buckthorn families) on its
alternate host European Buckthorn
Achene - one-seeded fruit in plants of the Aster (Composite) family (and others) that is often
accompanied by a pappus - Pappus - modified calyx in plants of the Aster (Composite) family that
is composed of bristles, hairs, teeth or scales that aid in seed dispersal - the picture at right
shows the achene and pappus of White Snakeroot (seed dispersed by wind) - the picture at
middle right shows the achene and pappus of Tall Beggar-ticks (seed dispersed by attaching to
mammals or birds via two barbed teeth known as awns) - the picture at far right shows the awns of
Ryegrass
Calyx - the outer segment of a flower that encloses a flower bud - the individual parts of the calyx
are petal-like structures called sepals - sepals are often green but when they are similar to (and
the same colour as) the petals they are referred to as tepals - the picture at right shows the calyx of
Hedge Bindweed - the picture at middle right shows the bladder-like calyx of Bladder Campion -
the picture at far right shows the sepals of Thyme-leaved Sandwort
Stamen, Staminate - see pistil
Spadix - a dense spike of small flowers on a fleshy stalk that is associated with (and sometimes
enclosed by) a spathe - the fruiting spadix is a dense cluster of berries - Spathe - a large bract -
characteristic of the Arum family - the picture at right shows the spadix and spathe of
Jack-in-the-Pulpit ("Jack" is the spadix, "Pulpit" is the spathe) - the picture at far right shows the
fruiting spadix of Jack-in-the-Pulpit
Composite - the Composite family (also known as Aster, Daisy or Sunflower family) is one of the
largest families of flowering plants - the flower head is not a single flower but a composite of many
disk and / or ray flowers (called florets or floscules) - the picture at top right shows Chicory which
only has ray florets - the picture at top far right shows White Snakeroot which only has disk florets -
the picture at bottom right shows Black-eyed Susan which has ray and disk florets - the picutre at
bottom far right shows a ray floret of Chicory containing both male and female parts - the ovary will
become an achene - 5 fused anthers form a tube around the style from which the stigma protrudes
- corolla consists of 5 fused petals
Corolla - collective term for all the petals of a flower - the corolla and calyx are together called the
perianth
Scutellum - In zoology, the shield-like plate or scale on an insect thorax - typically a small triangle
at the base of the wings pointing to the tip of the abdomen - In botany, the shield-like structure of a
grass seed - the picture at right shows the scutellum of a Transverse Hover Fly - the picture at far
right shows the large scutellum of a Green Stink Bug
Thorax - the mid-section of an insect to which the head (first or anterior section), abdomen (third or
posterior section), legs and wings are attached - composed of three segments, the prothorax (T1),
mesothorax (T2) and metathorax (T3) - Abdomen - the posterior section of an insect consisting of a
number of segments (11 in most orders but reduced in others) and containing the digestive tract
and reproductive organs - the picture at right shows the head, thorax and abdomen (segments A1
to A10) of a White Tail dragonfly - the picture at far right shows the head, thorax (segments T1 to
T3) and abdomen (segments A1 to A10) of an Underwing Moth larva
Haltere - also halter - modified wings of some insects that act as a gyroscope and help in
maintaining stability and executing high speed maneuvers during flight - in flies, the hindwings
have evolved into halteres while in some other species, the forewings have evolved - the picture at
right shows the halteres of a Band-winged Crane Fly - the picture at far right shows the halteres of
a Tachinid fly
Prothorax - the 1st or anterior (T1) of 3 segments in the thorax of an insect - bears the first pair of
legs - mesothorax - the 2nd (T2) of 3 segments in the thorax of an insect - bears the 2nd pair of
legs and the forewings (when present) - metathorax - the 3rd or posterior (T3) of 3 segments in the
thorax of an insect - bears the 3rd pair of legs and the hindwings (when present) - the picture at
right shows the forewings and hindwings of a Silvery Blue Butterfly
Pronotum - one of the main sclerites (exoskeletal plates) in the prothorax of an insect forming the
upper (dorsal) surface - in beetles and treehoppers, the pronotum is greatly exaggerated, aiding in
identification - the picture at right shows the pronotum of a Locust Borer Beetle - the picture at far
right shows the pronotum of a Buffalo Treehopper
Spiracle - respiratory pore, especially those located along the sides of insects - the spiracles of a
caterpillar (Butterflies and Moths) appear on the 1st thoracic segment (T1) and the 1st through 8th
abdominal segments (A1 to A8) - the coloured area circling some spiracles often aids in
identification - the picture at right shows the T1 spiracle of an Underwing Moth larva - the picture at
far right shows the A1 to A4 spiracles of a Pandorus Sphinx Moth larva
Stigma - in botany, see pistil - in zoology, a spiracle - also a coloured cell on the leading edge of
each wing in a dragonfly or damselfly and other insects (formerly pterostigma) - nodus - the
shallow notch at the midpoint of the leading edge of each wing in a dragonfly or damselfly - origin
of several veins that add strength and flexibility to the wing - the picture at right shows the stigma
and nodus of a 4-spotted Skimmer Dragonfly
Median stripe (damselflies) - dark stripe that runs along the centre of the thorax, sometimes
bisected by the pale Middorsal stripe - Antehumeral stripe - pale stripe below the median stripe on
either side of the thorax - Humeral stripe - dark stripe below the antehumeral stripe on either side
of the thorax - additional stripes may be present below the humeral stripe - pattern on dragonflies
is more complex - Postocular spots - pale marks just below the eyes (not always present) - the
picture at right shows the median stripe and postocular spot of an Eastern Forktail Damselfly - the
picture at far right shows the stripes of a Slender Spriead-wing Damselfly
Annual / Biennial / Perennial - life cycle duration of a plant - an annual plant germinates, flowers and
dies in one year - a biennial plant takes two years to complete its life cycle - in the 1st year, it
produces only leaves low to the ground, often in a rosette - remains dormant over the winter and
flowers in the 2nd year - a perennial plant lives more than two years - herbaceous refers to small
flowering plants (annual, biennial or perennial) that have no persistent woody stem - forb refers to a
herbaceous plant that is not a graminoid (grasses, sedges, rushes) - woody plants are perennial
only, usually referring to shrubs and trees - the picture at right shows 1st year Common Mullein
(biennial) - the picture at far right shows 2nd year Common Mullein
Bract - modified leaf that performs a specialized function such as attracting pollinators or
protecting young flowers - the picture at right shows the bracts of Field Bindweed - the picture at
middle right shows the bract (called a spathe in this case) of Skunk Cabbage - the picture at far
right shows the bracts of Wild Teasel
Monoecious - a plant that has male and female flowers (sometimes simultaneously, sometimes
at different times) - Dioecious - a plant that has only male (androecious - pollen / microspore
producers) or female (gynoecious - seed / megaspore producers) flowers - a hermaphrodite plant
(the majority of plants) has a flower with both male and female reproductive parts - the picture at
right shows the male and female flowers of Pennsylvania Sedge (monoecious) - the picture at
middle right shows the male and female flowers (on separate plants) of White Campion
(dioecious) - the picture at far right shows the pistil and stamens of Fringed Loosestrife
(hermaphrodite)
Simple Leaf - a leaf composed of only 1 blade attached to a petiole (petiolated) or directly to the
stem (sessile) - Compound Leaf - a leaf composed of 2 or more leaf blades (pinnae or leaflets)
attached to a rachis (pinnately compound) or the tip of a petiole (palmately compound) - a pinnately
compound leaf may be odd pinnate (leaflet at tip) or even pinnate - a bi-pinnately compound leaf is
one where each leaflet is also pinnately compound - a trifoliate leaf is a palmately compound leaf
that has 3 leaflets - the picture at right shows the simple leaf of Velvet-leaf - the picture at middle
right shows the odd pinnately compound leaf of Staghorn Sumac - the picture at far right shows the
palmately compound (trifoliate) leaf of Red Clover
Prolegs - also known as false legs or pseudopods, the fleshy stubs found on the abdomen of
most butterfly, moth and sawfly larva as well as some fly larva - used for locomotion but unlike true
legs on the thorax of the larva which will become the legs of the adult, the prolegs have no
segments / joints and no counterpart in the adult - with few exceptions, sawfly larvae have 6 or
more pairs of prolegs while moth and butterfly larvae have up to 5 pairs - the picture at right shows
the true legs and the prolegs of a Zebra Caterpillar Moth larva
Native - plant that is found in the area where it evolved - Non-native - plant introduced to an area by
humans, intentionally or accidentally - Invasive - non-native plant which easily establishes itself
and spreads quickly causing economic or environmental damage - Naturalized - non-native plant
that does not need human assistance to reproduce - Exotic - plant that is not native to the continent
on which it is found - Weed - native or non-native plant that is not wanted where it is growing and
may pose a threat to the environment or to agriculture
Dimorphism (sexual) - visual or behavioural difference between males and females of the same
species - male birds often have evolved ornamental plumage in competition over mates - female
birds however often have a drab appearance to provide camouflage while nesting - sometimes
these traits, being costly to produce or maintain, conspicuous or a hindrance in flight, are an
impediment to survival as in the Peacock at right - the picture at far right shows the visual
dimorphic traits of male and female Mallard Ducks
Rhizome - in botany, a modified stem - usually horizontal and underground - often sends out roots
and shoots from nodes - each piece may be able to produce a new plant
Symbiosis - a relationship between two or more organisms which can be one of four types:
Parasitism - a relationship where one organism is harmed and the other benefits -
Commensalism - a relationship where there is no harm or benefit to one organism but the other
benefits - Mutualism - a relationship where both organisms benefit - Amensalism - a relationship
where there is no harm or benefit to one organism but the other is harrmed - some say that
symbiosis only refers to mutualism - the picture at right shows the parasitic Dodder plant
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Alternate / Opposite - the arrangement of leaves or branches on a stem - alternate - each leaf (or
branch) arises from a different point on the stem on alternate sides - opposite - two leaves (or
branches) arise from the same point on the stem on opposite sides - an arrangement of leaves
may also be classified as whorled if several leaves originate (or appear to) from the same point on
a stem - the picture at right shows the alternate leaves of White Elm - the picture at middle right
shows the alternate branches of White Elm - the picture at far right shows the opposite branches
of European Euonymus - MAD is an acronym that can be used to describe most trees that have
oppostie branching (Maples, Ashes, Dogwoods)
Thallus - a plant body that is not differentiated into stems and leaves - lacks true roots and a
vascular system - algae, fungi, lichens and some liverworts have thalli - the picture at right shows
the thallus of Snake Liverwort
Vascular - in plants, the system of conducting tissues and supporting fibres - xylem tissue
transports water and minerals from the roots to the leaves and the rest of the plant - phloem tissue
transports food (photosynthate - products of photosynthesis) from the leaves to the rest of the plant
- xylem tissue dies each year (forming the rings in tree trunks from which the age can be
determined) - phloem tissue lives as long as the plant or tree - Mosses, Liverworts, Hornworts and
Algae lack a vascular system
Photosynthesis - using chlorophyll, the process by which plants, some bacteria and some
protistans (single-celled organisms) convert light energy into chemical energy - carbohydrates
(sugars for example) are synthesized from carbon dioxide and water producing oxygen in the
process - responsible for maintaining oxygen levels in the atmosphere and supplying most of the
energy needed for life
Coniferous - cone-bearing tree or shrub usually with needles or scales for leaves which they
retain year round - most coniferous trees are referred to as softwoods or evergreens - Deciduous -
a tree or shrub whose leaves fall off yearly - most deciduous trees are referred to as hardwoods or
broadleaf - a conifer usually refers to a coniferous tree but a few species (tamaracks, larches and
Dawn Redwood for example) are known as deciduous conifers as their needles fall off yearly - a
coniferous tree photsynthesizes (at a slower rate than the broad leaves of deciduous trees) year
round and their needles lose less water and are more wind-resistant allowing them to survive in
harsher climates and poorer soil conditions compared to deciduous trees
Median area - Moths - the central portion of the forewing - Basal (Antemedian or AM) area - the
area at the base of the forewing, often separated from the median area by the Antemedial (AM) line
- Postmedian (PM) area - the area in the outer third of the forewing, often bordered by the
Subterminal (ST) line and the Postmedial (PM) line - Terminal line - the outermost line on the
forewing - these parameters often aid in identifying noctuid moths (Owlet, Dart, Litter, Underwing) -
the picture at right shows these paramaters on a Black-banded Owlet Moth - the picture at far right
shows the ST line of a Pearly Wood Nymph Moth
Node - the positon on the stem where leaves, branches or flowers are attached - internode - the
region of the stem between nodes- sessile - leaf attached to the stem with no visible stalk -
petiolate - leaf attached to the stem by a stalk (petiole) - clasping - sessile leaf in which the base
partially or entirely surrounds the stem - perfoliate - sessile leaf in which the base is fused around
the stem and the stem appears to pierce the leaf - connate perfoliate - bases of opposite leaves
fused around the stem - sheathed - a tubular shaped portion of the leaf base surrounds the stem -
the pictures at right show the sessile leaf of Daisy Fleabane, the petiolate leaf of Velvet-leaf, the
clasping leaf of New England Aster, the perfoliate leaf of Bellwort, the connate perfoliate leaf of
Boneset and the sheathed leaf of Barnyard Grass
Inflorescence - cluster of flowers or groups of flowers - characterized by the arrangement of flowers
on the peduncle - solitary - only 1 flower on the peduncle - spike - group of sessile (no stalk)
flowers on the rachis - raceme - a spike but the flowers have pedicels - panicle - branched or
compound raceme where each branch has more than 1 flower - umbel - a flat-topped or rounded
raceme where individual flower stalks arise from a common center - characteristic of the Carrot
family - corymb - a raceme where individual flower stalks are of different lengths such that all
flowers are at the same level giving it a flat-topped appearance - head or capitulum - short dense
spike of flowers that appears to be a single flower - characteristic of the Aster family (see
composite) - cyme - a raceme in which the main axis and branches terminate in a single flower -
catkin or ament - a usually hanging spike, most of which are composed of unisexual flowers -
typically found on trees such as willows and cottonwoods - see also spadix for another type of
inflorescence - the pictures at right show the spike of Narrow-leaved Cattail, the raceme of
Wild Lupine, the panicle of False Solomon's Seal, the umbel of Flowering-rush, the cyme of
St. John's Wort and the catkins of Ironwood
Connate Perfoliate - see Node
Axil - upper angle between a plant stem and a leaf, branch or petiole - an axillary bud develops in
the axil of a leaf
Elytra (singular Elytron) - modified, hardened forewings of certain insect orders, especially beetles
(Coleoptera) - primarily serve as protection for the more delicate hindwings which are used for
flying - in some ground beetles (Carabidae) and others the elytra are fused together making flight
impossible - the picture at right shows the elytra of the Spotted Cucumber Beetle
Metamorphosis - some insects, fish, amphibians, crustaceans and others undergo
metamorphosis, a process where an animal changes form during development (a tadpole
becomes a frog, a caterpillar becomes a butterflly) - some insects undergo complete
metamorphosis (egg to larva to pupa to imago / adult) in which the larva enters the nonfeeding
pupa stage (protective cocoon or hardened case) and undergoes a complete transformation - a
butterfly pupa is called a chrysalis - other insects undergo incomplete metamorphosis (egg to
nymph to imago / adult) where the nymph (usually resembling the adult without wings) goes
through stages of growth and moulting to become an adult - these stages are called instars
(larvae also go through these stages) - an aquatic nymph (dragonfly, stonefly and others) is called
a naiad - the top picture at right shows a Monarch larva - the top picture at centre shows a Monarch
butterfly (complete metamorphosis) - the top picture at far right shows a recently moulted Cicada
imago / adult (incomplete metamorphosis) resting on its exuvia - the bottom picture at right shows
a Baltimore Checkerspot chrysalis (pupa) - the bottom picture at far right shows a damselfly naiad
Pistil - the female part of a flower consists of the ovary (will become the fruit) and the stalk-like style
which supports the pollen receiving stigma - the ovary contains1 or more ovules which will
become the seeds after fertilization - Stamen - the male part of a flower consists of the stalk-like
filament which supports the pollen producing anther - the number, appearance and arrangement
of pistils and stamens varies - some flowers have only a pistil (carpellate or pistillate) or only a
stamen (staminate) - the picture at right shows the reproductive structure of Enchanter's
Nightshade - the picture at middle right shows the anthers and stigmas of Chicory - the picture at
far right shows the anthers and stigmas of Switch Grass
Slime Mould - group of amoeba-like single cell organisms which group together and fuse
exhibiting characteristics of fungi, plants and animals - there are 2 stages to a slime mould's life
cycle - the first (plasmodial) allows for movement and feeding - under adverse conditions (cold,
lack of food) the mould enters a second, immobile stage (sporangial) where it fruits and
reproduces via spores - the picture at right shows the sporangial stage of Chocolate Tube Slime
Mould - the middle and far right pictures show the plasmodial and sporangial stages of False
Puffball Slime Mould
Peduncle - stalk bearing an inflorescence or a solitary flower (or fruit) on the main stem - Rachis -
central axis bearing a flower or flowers in an inflorescence (spike, raceme or panicle) or a pinna in
a fern blade or pinnately compound leaf - extension of the peduncle in the case of flowers, the
petiole in the case of a compound leaf or the stipe in the case of a fern - Pedicel - stalk bearing a
single flower in an inflorescence - Petiole - stalk that joins a simple or compound leaf blade to the
stem - Pinna (plural pinnae) - leaflet of a fern frond or a pinnately compound leaf - a leaf has a bud,
a pinna does not - Petiolule - stalk that joins a pinna to the rachis - Pinnule or Pinnula - leaflet of a
pinna in a fern frond or a bi-pinnately compound leaf - Stipe - stalk that joins a fern blade to the
stem - the picture at top right shows the peduncle of Spotted Water Hemlock - the picture at top
middle right shows the pedicel and rachis of an Enchanter's Nightshade raceme - the picture at
top far right shows the rachis, pinna and petiole of a pinnately compound Mountain Ash leaf - the
picture at bottom right shows the rachis, stipe and pinna of a Bracken Fern frond - the picture at
bottom far right shows a close-up of 3 pinnae of Bracken Fern
Sorus (plural sori) - cluster of sporangia (structures producing and containing spores) used in
reproduction for ferns, fungi and lichens - on ferns sori occur on fertile fronds only and they turn
from green to brown as they mature - for sporangia, see also slime mould - the picture at right
shows the sori of Bulblet Fern - the picture at far right shows the sori of Marginal Wood Fern
Melanistic - an excessive production of melanin (skin or hair pigment) resulting in a dark colour -
Amelanistic - a lack of melanin (albino) - the picture at right shows a melanistic Garter Snake - the
picture at far right shows an amelanistic Green Frog
Antenna (plural antennae) - jointed sensory appendages attached to the head of arthropods - most
arthropods have 1 pair of antennae (spiders have none, crustaceans have 2 pairs) - most notably
used to provide a sense of smell but may also provide information on touch, sound (vibration),
humidity, air motion or heat - consists of 3 parts: Scape (basal segment controls movement),
Pedicel (controls movement and contains sense organs) and Flagellum (1 to more than 30
structures known as flagellomeres containing sense organs) - the scape, pedicel and
flagellomeres are collectively known as antennomeres - in bees, the male has 11 flagellomeres
and the female has 10 - the picture at right shows the antennae of a Common Carpenter Bee
Flagellum, Flagellomere - see antenna
Parthenogenesis - asexual reproduction where the growth and development of an embryo
occurs without fertilization
Instar - stage in the life of the larva, nymph or pupa of arthropods where they moult to shed their
exoskeleton allowing them to grow - the period between each moult is known as an instar -
successive instars are often characterized by a different form, pattern, colour or number of
segments - the number of instars for a given organism depends on the species and
environmental conditions - exuvia - remains of the exoskeleton that is left after moulting - the
picture at right shows an early instar of a Spicebush Swallowtail larva - the picture at far right
shows a Cicada nymph
Lichen - mutualistic association of a fungus and a photosynthesising partner (alga or
cyanobacterium) - the fungal component may not be able to survive on its own - the Lichen takes
its binomial name (genus:species) from that of the fungus - most of the fungal components
belong to the Sac fungi (Ascomycota) division - many lichens are sensitive to pollution and will not
be present where air quality is poor - the picture at right shows a Physcia species lichen
Drake - male duck
Eclipse - Dull coloured female-like plumage of male ducks (drakes) into which they moult after
breeding - for a brief period they are unable to fly as the flight feathers are lost and renewed - some
drakes remain in eclipse for a month or two while others retain their eclipse plumage until spring
when there is a further moult to enter breeding plumage - the flight feathers are not affected during
this moult - most non-waterfowl birds lose their flight feathers sequentially during a moult so that
they retain the ability to fly
Reniform spot - usually kidney-shaped spot or outline in the outer median area of a moth's
forewing - Subreniform spot - small spot or outline between the Reniform spot and the inner edge
of a moth's forewing - Orbicular spot - round spot or outline in the inner median area of a moth's
forewing - these parameters often aid in identifying noctuid moths (Owlet, Dart, Litter, Underwing) -
the picutre at right shows the Reniform and Subreniform spots of an Ilia Underwing moth - the
picture at far right shows the Reniform and Orbicular spots of a Grote's Sallow moth
Teneral - a teneral insect is one that has recently moulted or emerged from the pupa and has a
soft exoskeleton and pale colouration - it may be able to fly with difficulty but until its exoskeleton
and wings have hardened it is in a vulnerable state
Eclosure / Eclosion - in biology, the emergence of an adult insect from the pupa or a larva from an
egg - verb eclose
Areolet - For insects, a small area on the wings enclosed by veins - the picture at right shows the
aerolets of an Ophion Ichneumon Wasp
Perigynium (plural perigynia) - in sedges, a sac-like structure that encloses the achene - the
picture at right shows the perigynia of Bladder Sedge